Actually, if you were going to have the swirl tank under suction (as well as the radiator), as recommended by Carroll Smith - I would think they would still auto-bleed.
As you probably know, the bottom of the expansion tank is "sucked" by the water pump, and the swirl, radiator and head bleeds (if you have them) go to the top of the expansion tank. George has a good diagram on his cooling system sticky.
If the expansion tank top was lower than the radiator, though - I would guess you would figure it our pretty quick!
Here's the picture...
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